One Galápagos tortoise lived 141 years, turning quiet meals and slow steps into a legacy of care, memory, and conservation ...

LOS ANGELES (AP) — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has ...

No one knows exactly when Gramma was born. But if the estimated birth year of 1884 is accurate, Chester Arthur occupied the Oval Office and there were only 39 states at the time.

Giant tortoises are some of the long-lived species on the planet. Here’s what biologists now understand about one of nature’s ...